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In Drosophila, genes a and b are located at positions 22.0 and 42.0 on chromosome 2, and genes c and d are located at positions 10.0 and 25.0 on chromosome 3. A fly homozygous for the wild-type alleles of these four genes was crossed with a fly homozygous for the recessive alleles, and the F1 daughters were backcrossed to their quadruply recessive fathers. What offspring genotypes would you expect from this backcross, and in what proportions?

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