Using bacteriophage P22, you performed a three-factor cross in Salmonella typhimurium. The cross was between an Arg-Leu-His- recipient bacterium and bacteriophage P22, which was grown on an Arg+ Leu+ His+ strain. You selected for 1000 Arg+ transductants and tested them on several selective media by replica plating. You obtained the following results:
Arg+Leu-His- 585 Arg+Leu-His+ 300
Arg+Leu+His+ 114 Arg+Leu+His- 1
What is the order of the three markers?